Browsing by Funding "Acknowledgements – We acknowledge Russell Lande for discussions of the results and Kate Layton Matthews for comments on the manuscript. Funding – This work was supported by the European Economic Area (EEA) Grants UCM-EEA-ABEL-02-2009 and 005-ABEL-CM-2014A under the NILS – Science and Sustainability Programme, by the SUSTAIN project of the Research Council of Norway and by the Research Council of Norway (SFF-III 223257/F50). JJ acknowledge financial support of Ministerio de Educación, Cultura y Deporte through grant FPU13/02934. FJC acknowledge financial support through grants FIS2010-17440 and FIS2006-05895 of Ministerio de Ciencia e Innovación (Spain), FIS2015-67745-R (MINECO/FEDER) of Ministerio de Economía y Competitividad (Spain) and Fondo Europeo de Desarrollo Regional (FEDER, EU), GR35/14-920911, GR35/10-A-920911 and GR58/08-920911 of Universidad Complutense de Madrid and Banco Santander (Spain). Author contributions – SE and FJC designed the research"," JJ and FJC performed the research"," JJ, BE, SE and FJC wrote the manuscript. Competing interests – We have no competing interests."